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Journal Articles
McClelland, C.J.R., Denny, C.K., Larsen, T.A., Stenhouse, G.B. & Nielsen, S.E. 2021. Landscape estimates of carrying capacity for grizzly bears using nutritional energy supply for management and conservation planning. Journal of Nature Conservation 62: 126018.
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DOI: https://doi.org/10.1016/j.jnc.2021.126018
Tracy S. Lee. T.S., Creech, T.G., Martinson, A., Nielsen, S.E., Jakes, A.F., Jones, P.F., Sanderson, K. & Ford, A.T. 2021. Prioritizing human safety and multispecies connectivity across a regional road network. Conservation Science & Practice 3(2): e327.
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DOI: 10.1111/csp2.327
Lamb, C.L., Ford, A.T., McLellan, B.N., Proctor, M.F., Mowat, G., Ciarniello, L., Nielsen, S.E. & Boutin, S. 2020. The ecology of human–carnivore coexistence. Proceedings of the National Academy of Sciences 117(30): 17876-17883.
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DOI: 10.1073/pnas.1922097117
McClelland, C.J.R., Coops, N.C., Kearney, S.P., Burton, A.C., Nielsen, S.E. & Stenhouse, G.B. 2020. Variations in grizzly bear habitat selection in relation to the daily and seasonal availability of annual plant-food resources. Ecological Informatics 58: 101116.
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DOI: 10.1016/j.ecoinf.2020.101116
McClelland, C.J.R.,  Coops, N.C., Berman, E.E., Kearney, S.P., Nielsen, S.E., Burton, A.C. & Stenhouse, G.B. 2020. Detecting changes in understory and canopy vegetation cycles in West Central Alberta using a fusion of Landsat and MODIS. Applied Vegetation Science 23(2), 223-238.
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DOI: 10.1111/avsc.12466
Souliere, C.M., Coogan, S.C.P., Stenhouse, G.B. & Nielsen, S.E. 2020. Harvested forests as a surrogate to wildfires in relation to grizzly bear food-supply in west-central Alberta. Forest Ecology and Management Volume 456 (117685), 1-13.
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DOI: 10.1016/j.foreco.2019.117685
Pollock, S.Z., Whittington, J., Nielsen, S.E. & St. Clair, C.C. 2019. Spatiotemporal railway use by grizzly bears in Canada's Rocky Mountains. Journal of Wildlife Management 83(8), 1787-1799.
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DOI: 10.1002/jwmg.21750
Kearney, S.P., Coops, N.C., Stenhouse, G.B., Nielsen, S.E., Hermosillae T., White, J.C. & Wulder, M.A. 2019. Grizzly bear selection of recently harvested forests is dependent on forest recovery rate and landscape composition. Forest Ecology and Management 449(1):117459.
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DOI: 10.1016/j.foreco.2019.117459
Laskin, D.N., McDermid, G.J., Nielsen, S.E., Marshall, S.J., Roberts, D.R. & Montaghi, A. 2019. Advances in phenology are conserved across scale in present and future climates. Nature Climate Change 9:419–425.
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DOI: 10.1038/s41558-019-0454-4
Gilhooly, P.S., Nielsen, S.E., Whittington, J. & Cassady St. Clair, C. 2019. Wildlife mortality on roads and railways following highway mitigation. Ecosphere 10(2): e02597.
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DOI: 0.1002/ecs2.2597
Hughes, C. & Nielsen, S.E. 2019. ‘Bear are only the Lightning Rod’: Ongoing Acrimony in Alberta’s Grizzly Bear Recovery. Society & Natural Resources: An International Journal  32(1):34-52.
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DOI: 10.1080/08941920.2018.1502853
Ladle, A., Avgar T., Wheatley, M., Stenhouse, G.B., Nielsen, S.E. & Boyce, M.S. 2019. Grizzly bear response to spatio-temporal variability in human recreational activity. Journal of Applied Ecology 56:375–386.
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DOI: 10.1111/1365-2664.13277
Larsen, T.A., Nielsen, S.E., Cranston, J. & Stenhouse, G.B. 2019. Do remnant retention patches and forest edges increase grizzly bear food supply? Forest Ecology and Management 433:741-761.
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DOI: 10.1016/j.foreco.2018.11.031
Coogan, S.C.P., Coops, N.C., Janz, D.M., Cattet, M.R.L., Kearney, S.P., Stenhouse, G.B. & Nielsen, S.E. 2019. Towards grizzly bear population recovery in a modern landscape. Journal of Applied Ecology 56(1):93-99.
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DOI: 10.1111/1365-2664.13259
Denny, C.K., Stenhouse, G.B & Nielsen, S.E. 2018. Scales of selection and perception: landscape heterogeneity of an important food resource influences habitat use by a large omnivore. Wildlife Biology 2018(1):wlb.00409.
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DOI: 10.2981/wlb.00409
Boulanger, J., Nielsen, S.E. & Stenhouse, G.B. 2018. Using spatial mark-recapture for conservation monitoring of grizzly bear populations in Alberta. Scientific Reports 9:5204.
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DOI: 10.1038/s41598-018-23502-
Lamb, C.T., Mowat, G., Reid, A., Smit, L., Proctor, M., McLellan, B.N., Nielsen, S.E., Boutin, S. 2018. Effects of habitat quality and access management on the density of a recoverying grizzly bear population. Journal of Applied Ecology 55:1406–1417.
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DOI: 10.1111/1365-2664.13056
Pollock, S.Z., Nielsen, S.E., & St. Clair, C.C. 2017. A railway increases the abundance and accelerates the phenology of bear-attracting plants in a forested, mountain park. Ecosphere 8(10):e01985. 10.1002/ecs2.1985.
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DOI: 10.1002/ecs2.1985
Lamb, C.T., Mowat, G., Gilbert, S.L., McLellan, B.N., Nielsen, S.E. & Boutin S. 2017. Density-dependent signaling: An alternative hypothesis on the function of chemical signaling in a non-territorial solitary carnivore. PLoS ONE 12(10): e0184176.
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DOI: 10.1371/journal.pone.0184176
Nielsen, S.E., Larsen, T.A., Stenhouse, G.B. & Coogan, S.C.P. 2017. Complementary food resources of carnivory and frugivory affect local abundance of an omnivorous carnivore. Oikos 126: 369–380.
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DOI: 10.1111/oik.03144
Lamb, C.T., Mowat, G., McLellan, B.N., Nielsen, S.E. & Boutin, S. 2017. Forbidden fruit: human settlement and abundant fruit create an ecological trap for an apex omnivore. Journal of Animal Ecology 86(1): 55–65.
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DOI: 10.1111/1365-2656.12589
Denny, C.K. & Nielsen, S.E. 2017. Spatial heterogeneity of the forest canopy scales with the heterogeneity of an understory shrub based on fractal analysis. Forests 8(5): 146.
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DOI: 10.3390/f8050146
Laskin, D.N., Montaghi A, Nielsen, S.E. & McDermid, G.J. 2016. Estimating understory temperatures using MODIS LST in mixed cordilleran forests. Remote Sensing 8: 658.
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DOI: 10.3390/rs8080658
Cristescu, B., Stenhouse, G.B., Symbaluk, M., Nielsen, S.E. & Boyce, M.S. 2016. Wildlife habitat selection on landscapes with industrial disturbance. Environmental Conservation 1-10.
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DOI: 10.1017/S0376892916000217
Braid, A.C.R., Manzer, D. & Nielsen, S.E. 2016. Wildlife habitat enhancements for grizzly bears: Survival rates of planted fruiting shrubs in forest harvests. Forest Ecology and Management 369: 144-154.
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DOI: 10.1016/j.foreco.2016.03.032
Cristescu, B., Stenhouse, G.B., Symbaluk, M., Nielsen, S.E. & Boyce, M.S. 2016. Wildlife habitat selection on landscapes with industrial disturbance. Environmental Conservation [in press]
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DOI: 10.1017/S0376892916000217
Braid, A.C.R. & Nielsen, S.E. 2015. Prioritizing sites for protection and restoration for grizzly bears (Ursus arctos) in southwestern Alberta, Canada. PLoS ONE 10(7):e0132501.
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DOI: 10.1371/journal.pone.0132501
Rovang, S., Nielsen, S.E. & Stenhouse, G. 2015. In the trap: detectability of fixed hair trap DNA methods in grizzly bear population monitoring. Wildlife Biology 21(2): 68–79.
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DOI: 10.2981/wlb.00033
Nijland, W., Coops, N.C., Nielsen, S.E. & Stenhouse, G. 2015. Integrating optical satellite data and airborne laser scanning in habitat classification for wildlife management. International Journal of Applied Earth Observation and Geoinformation 38: 242–250.
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DOI: 10.1016/j.jag.2014.12.004
Nijland W., Nielsen S.E., Coops N.C., Wulder M.A. & Stenhouse G.B. 2014. Fine-spatial scale predictions of understory species using climate-and LiDAR-derived terrain and canopy metrics. Journal of Applied Remote Sensing 8(1): 083572-083572.
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DOI: 10.1117/1.JRS.8.083572
Coogan S.C.P., Raubenheimer D., Stenhouse G.B. & Nielsen S.E. 2014. Macronutrient optimization and seasonal diet mixing in a large omnivore, the grizzly bear: A geometric analysis. PloS ONE 9(5): e97968.
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DOI: 10.1117/1.JRS.8.083572
Pigeon K.E., Nielsen S.E., Stenhouse G.B. & Côté S.D. 2014. Den selection by grizzly bears on a managed landscape. Journal of Mammalogy 95(3): 559-571.
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DOI: 10.1644/13-MAMM-A-137
Roberts D.R., Nielsen S.E. & Stenhouse G.B. 2014. Idiosyncratic responses of grizzly bear habitat to climate change based on projected changes in their food resources. Ecological Applications 24: 1144-1154.
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DOI: 10.1890/13-0829.1
Boulanger J., Cattet M., Nielsen S.E., Stenhouse G. & Cranston J. 2013. Use of multi-state models to explore relationships between changes in body condition, habitat and survival of grizzly bears Ursus arctos horribilis. Wildlife Biology 19: 274-288. 
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DOI: 10.2981/12-088
Nielsen S.E., Cattet M.R.L., Boulanger J., Cranston J., McDermid G.J., Shafer A.B.A., Stenhouse G.B. 2013. Environmental, biological and anthropogenic effects on grizzly bear body size: temporal and spatial considerations. BMC Ecology 13: 31.
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DOI: 10.1186/1472-6785-13-31
López-Alfaroa C., Robbins C.T., Zedrosser A. & Nielsen S.E. 2013. Energetics of hibernation and reproductive trade-offs in brown bears. Ecological Modeling 270: 1-10.
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DOI: 10.1016/j.ecolmodel.2013.09.002
Stewart B.P., Nelson T.A., Laberee K., Nielsen S.E., Wulder M.A. & Stenhouse G. 2013. Quantifying grizzly bear selection of natural and anthropogenic edges. Journal of Wildlife Management 77(5): 957-964.
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DOI: 10.1002/jwmg.535
Nijland W., Coops N.C., Coogan S.C.P., Bater C.W., Wulder M.A., Nielsen S.E., McDermid G. & Stenhouse G.B. 2013. Vegetation phenology can be captured with digital repeat photography and linked to variability of root nutrition in Hedysarum alpinum. Applied Vegetation Science 16(2): 317–324.
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DOI: 10.1111/avsc.12000
Girard T.L., Bork E.W., Nielsen S.E. & Alexander M.J. 2013. Seasonal variation in habitat selection by free-ranging feral horses within Alberta's Forest Reserve. Rangeland Ecology & Management 66(4): 428-437.
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DOI: 10.2111/REM-D-12-00081.1
Girard T.L., Bork E.W., Nielsen S.E. & Alexander M.J. 2013. Landscape-scale factors affecting feral horse habitat use during summer within the Rocky Mountain foothills. Environmental Management 51(2): 435-447.
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DOI: 10.1007/s00267-012-9987-2
Smulders M., Nelson T.A., Jelinski D.E., Nielsen S.E., Stenhouse G.B. & Laberee K. 2012. Quantifying spatial–temporal patterns in wildlife ranges using STAMP: A grizzly bear example.   Applied Geography 35(1–2): 124–131.
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DOI: 10.1016/j.apgeog.2012.06.009
Stewart B.P., Nelson T.A., Wulder M.A., Nielsen S.E. & Stenhouse G. 2012. Impact of disturbance characteristics and age on grizzly bear habitat selection. Applied Geography 34: 614-625.
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DOI: 10.1016/j.apgeog.2012.03.001
Coops N.C., Hilker T., Bater C.W., Wulder M.A., Nielsen S.E., McDermid G. & Stenhouse G. 2012. Linking ground-based to satellite derived phenological metrics in support of habitat assessment. Remote Sensing Letters 3(3): 191-200.
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DOI: 10.1080/01431161.2010.550330
Serrouya R., McLellan B.N., Boutin S., Seip D.R. & Nielsen S.E. 2011. Developing a population target for an overabundant ungulate for ecosystem restoration. Journal of Applied Ecology doi: 10.1111/j.1365-2664.2011.01998.x.
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DOI: 10.1111/j.1365-2664.2011.01998.x
Bater C.W., Coops N.C., Wulder M.A., Nielsen S.E., McDermid G. & Stenhouse G.B. 2011. Design and installation of a camera network across an elevation gradient for habitat assessment. Instrumentation Science & Technology 39: 231-247.
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DOI: 10.1080/10739149.2011.564700
Nielsen S.E., McDermid G., Stenhouse G.B. & Boyce M.S. 2010. Dynamic wildlife habitat models: Seasonal foods and mortality risk predict occupancy-abundance and habitat selection in grizzly bears. Biological Conservation 143: 1623-1634.
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DOI: 10.1016/j.biocon.2010.04.007
Nielsen S.E., Cranston J. & Stenhouse G.B. 2009. Identification of priority areas for grizzly bear conservation and recovery in Alberta, Canada. Journal of Conservation Planning 5: 38-60.
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Nielsen S.E., Boyce M.S., Beyer H., Huettmann F. & Stenhouse G.S. 2008. Can natural disturbance-based forestry rescue a declining population of grizzly bears? Biological Conservation 141: 2193-2207.
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DOI: 10.1016/j.biocon.2008.06.020
Nielsen S.E., Boyce M.S. & Stenhouse G.B. 2006. A habitat-based framework for grizzly bear conservation in Alberta. Biological Conservation 130: 217-229.
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DOI: 10.1016/j.biocon.2005.12.016
Munro R.H.M., Nielsen S.E., Price M.H., Stenhouse G.B. & Boyce M.S. 2006. Seasonal and diel patterns of grizzly bear diet and activity in west-central Alberta. Journal of Mammalogy 87: 1112-1121.
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DOI: 10.1644/05-MAMM-A-410R3.1
Nielsen S.E., Herrero S., Boyce M.S., Benn B., Mace R.D., Gibeau M.L. & Jevons S. 2004. Modelling the spatial distribution of human-caused grizzly bear mortalities in the Central Rockies Ecosystem of Canada. Biological Conservation 120: 101-113.
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DOI: 10.1016/j.biocon.2004.02.020
Nielsen S.E., Boyce M.S., Stenhouse G.B. & Munro R.H.M. 2003. Development and testing of phenologically driven grizzly bear habitat models. Ecoscience 10: 1-10.
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Nielsen S.E., Boyce M.S., Stenhouse G.B. & Munro R.H.M. 2002. Modeling grizzly bear habitats in the Yellowhead Ecosystem of Alberta: Taking autocorrelation seriously. Ursus 13: 45-56.
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Book Chapters
Research Reports
Thesis
Lamb, C.T. 2019. Grizzly bear population dynamics across productivity and human influence gradients. Ph.D. Thesis, University of Alberta, Edmonton, Alberta, Canada.
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Hughes, C. 2018. The importance and influence of the human dimensions in grizzly bear (Ursus arctos) conservation. Ph.D. Thesis, University of Alberta, Edmonton, Alberta, Canada.
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McLellan, C.R. 2018. Food availability and grizzly bear (Ursus arctos) selection of post-fire and thinned forests in the mountain national parks of Canada. M.Sc. Thesis, University of Alberta, Edmonton, Alberta, Canada.
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Braid, A.C.R. 2015. Mitigating the effects of human activity on grizzly bears (Ursus arctos) in southwestern Alberta. M.Sc. Thesis, University of Alberta, Edmonton, Alberta, Canada.
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DOI: 10.7939/R3ZD0P
Nielsen S.E. 2005. Habitat ecology, conservation, and projected population viability of grizzly bears (Ursus arctos L.) in west-central Alberta, Canada. PhD Dissertation, University of Alberta, Edmonton, Alberta, Canada.
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Outreach